Peptide Bioregulator Research Library: Publications from the Khavinson Program

By Vita Stream Inc · Updated October 1, 2026 · 21 publications and 2 books · Links checked September 2026

This library collects the main publications behind the peptide bioregulator concept — the research program that began with Vladimir Khavinson's group in Leningrad in the 1970s and continues today. For each paper we note what was studied, on what model or population, by what route, and what was measured, and we link to the journal where it appeared. We do not summarize the authors' conclusions: those belong in the papers, with their methods and limitations, and the links take you there.

If you are new to the terminology, start with What Are Bioregulators? Peptide Bioregulators and Khavinson Peptides, Explained Simply. This page assumes you know the difference between a natural tissue extract and a synthetic short peptide.

How this library is organized

The publications fall into five groups, and it helps to know which one you are reading before you open it.

Human research. Clinical reports on the original injectable preparations — Thymalin and Epithalamin — in defined groups of older adults. The oldest and longest studies in the program.

Animal research. Lifespan and hormone experiments with the synthetic tetrapeptide AEDG in fruit flies, rats, and monkeys, from 2000–2003.

Laboratory research. Cell-culture experiments, cytogenetics, and analytical chemistry with synthetic short peptides and with the natural extract itself.

Molecular modeling. Computer simulations of how short peptides could bind DNA or pass through transporter proteins. Theoretical by nature.

Review literature. Articles and books in which the researchers summarize their own program, plus one systematic review.

Every entry below carries one of these labels. A label tells you what kind of question the study could answer — and, just as importantly, what kind it could not.

Five things to check when you open a study

These are the questions we asked while annotating each entry. They work for any paper in this field.

A reading checklist for peptide bioregulator studies
Check	The question	Why it matters
Identity	Exactly which material was studied — a named injectable extract, a defined synthetic sequence, or something else?	Similar names (Epithalamin, Epitalon, Thymalin) refer to different substances.
Setting and route	Was it given to people or animals, and how — by injection, by mouth — or added to cells in a dish?	A cell culture is an experimental setting, not a way of taking something.
Comparison	Was there a control group, random allocation, blinding? For lab work, suitable controls?	Without a comparison, a change cannot be attributed to the intervention.
Endpoint	What was actually measured — a molecular marker, a questionnaire score, a clinical event?	These are different kinds of evidence and are not interchangeable.
Limits	How long, how many participants, which population, who funded it, and is there a correction or retraction?	The authors' own limitations section is usually the most useful paragraph in the paper.

Timeline of the program

Five decades of publications, at a glance
Years	What was published	Material	Note
1970s	Thymalin, Epithalamin isolated	Tissue extracts	Pharmaceutical development in Leningrad; the starting point of the program
2000–2003	Lifespan and hormone studies in flies, rats, monkeys; first cell studies	AEDG	The synthetic peptide is tested across models
2003	Human follow-up report	Thymalin, Epithalamin	266 participants, 6–8 years
2006 / 2011	Twelve- and fifteen-year cohort reports	Epithalamin	Same participants, two reports
2009–2013	Program reviews	Multiple	Narrative summaries by the originating group
2011–2019	Cell localization, chromatin, telomere studies	AEDG, EDR, KEDG	Laboratory work on mechanism
2016–2023	Modeling: DNA docking, structural motifs, transporters	Short peptides	Computational studies
2017	AEDG identified in the natural pineal complex	Natural extract	Analytical link between extract and synthetic sequence
2019–2020	Stem-cell culture studies	AEDG and other short peptides	Italian–Russian collaboration
2021	Systematic review	Literature	First review with a stated search method
2025	Cell-line study	AEDG	Outside group; published with a later correction

Human research

Three papers describe studies in people. All of them concern injectable preparations given in clinical settings by the originating group; none studied an oral product. Two report on the same participants at twelve and fifteen years.

Human research · 2003

Thymalin and Epithalamin: a six-to-eight-year follow-up report

Khavinson VKh, Morozov VG. Neuro Endocrinology Letters. 2003;24(3–4):233–240.

What was studied: Two of the original injectable preparations developed by the program — the thymus extract Thymalin and the pineal extract Epithalamin.
Model and setting: 266 older participants, followed for six to eight years. A historical clinical report from the originating group.
What was measured: Survival over the follow-up period and a set of physiological and clinical measures.
Where to read it: Publisher abstract · PubMed

Only the abstract is openly available; the administration schedule and methods are in the paper itself.

Human research · 2006

Epithalamin: a twelve-year clinical study in older adults

Korkushko OV, Khavinson VKh, Shatilo VB, Antonyuk-Shcheglova IA. Bulletin of Experimental Biology and Medicine. 2006;142:356–359.

What was studied: Epithalamin (spelled “epithalamine” in this paper), the pineal extract preparation, given by injection alongside the participants' usual care.
Model and setting: Older adults with signs of accelerated aging, enrolled through the Institute of Gerontology in Kyiv; twelve-year duration; described by the authors as randomized.
What was measured: Measures of physical work capacity and cardiovascular function, and survival over the follow-up period.
Where to read it: Publisher / DOI

A specific clinical population, not a general wellness population.

Human research · 2011

Epithalamin: fifteen-year follow-up of the same cohort

Korkushko OV, Khavinson VKh, Shatilo VB, Antonyk-Sheglova IA. Bulletin of Experimental Biology and Medicine. 2011;151:366–369.

What was studied: Epithalamin, given as six courses over three years in addition to usual care. Continues the cohort of the 2006 report.
Model and setting: 79 older adults — 39 receiving the peptide courses, 40 receiving usual care only — followed for fifteen years; described by the authors as a randomized comparative study.
What was measured: Age-related measures of the cardiovascular system, physical work capacity, the daily pattern of melatonin, metabolic markers, and survival in the two groups.
Where to read it: Publisher / DOI

Read together with the 2006 report; it is the same group of participants at a later time point.

Animal research

Three papers describe experiments with the synthetic tetrapeptide AEDG in animals, from fruit flies to rhesus monkeys. They belong to the early 2000s, when the synthetic peptide was first being tested across models. Animal studies answer questions about the animal model; they are a step in a research program, not a statement about people.

Animal research · 2000

AEDG in a fruit-fly lifespan experiment

Khavinson VKh, Izmaylov DM, Obukhova LK, Malinin VV. Mechanisms of Ageing and Development. 2000;120:141–149.

What was studied: The synthetic tetrapeptide AEDG (Ala-Glu-Asp-Gly), added to the nutrient medium.
Model and setting: Drosophila melanogaster (fruit flies), a standard invertebrate model for lifespan experiments.
What was measured: Lifespan of the fly populations under the tested conditions.
Where to read it: Publisher / DOI · PubMed

An invertebrate model; the earliest lifespan experiment with the synthetic peptide.

Animal research · 2001

AEDG and hormone measurements in old rhesus monkeys

Khavinson V, Goncharova N, Lapin B. Neuro Endocrinology Letters. 2001;22(4):251–254.

What was studied: The tetrapeptide AEDG, given by injection.
Model and setting: Old rhesus monkeys (Macaca mulatta) compared with young animals; a primate model from the Sukhumi/Adler primate center.
What was measured: Blood levels of melatonin and cortisol and their day–night pattern.
Where to read it: PubMed

A small primate study; a companion paper the same year reported melatonin and cortisol production in old monkeys.

Animal research · 2003

AEDG and melatonin secretion of the pineal gland in young and old rats

Kossoy G, Zandbank J, Tendler E, Anisimov V, Khavinson V, Popovich I, Zabezhinski M, Zusman I, Ben-Hur H. Journal of Endocrinological Investigation. 2003.

What was studied: The synthetic tetrapeptide AEDG.
Model and setting: Pineal glands of young and old rats; a laboratory study with an Israeli co-author group.
What was measured: Melatonin secretion by the pineal gland under the tested conditions.
Where to read it: Publisher / DOI · PubMed

Bridges the animal and laboratory groups: an organ-level measurement in rats of two ages.

Laboratory research

Eight papers describe work with short peptides in cell culture or in analytical chemistry. They concern molecules and cells under controlled conditions, not people. One of them (2017) asks a question that sits at the heart of the pillar article: whether the synthetic sequence AEDG is actually present in the natural pineal extract.

Cell culture · 2003

AEDG in cultured human fibroblasts

Khavinson VKh, Bondarev IE, Butyugov AA. Bulletin of Experimental Biology and Medicine. 2003;135:590–592.

What was studied: The synthetic tetrapeptide AEDG (Ala-Glu-Asp-Gly).
Model and setting: Cultured human fetal fibroblasts; the peptide was added directly to the culture medium.
What was measured: Telomerase activity and telomere length in the cultured cells.
Where to read it: Publisher / DOI · PubMed

A cell-culture experiment: it concerns cells in a dish, not people taking anything.

Cell culture · 2003

AEDG and chromatin in lymphocytes from elderly donors

Khavinson VKh, Lezhava TA, Monaselidze JR, Jokhadze TA, Dvalishvili NA, Bablishvili NK, Trofimova SV. Neuro Endocrinology Letters. 2003;24(5):329–333.

What was studied: The tetrapeptide AEDG, added to cell cultures.
Model and setting: Cultured blood lymphocytes from elderly donors; a Georgian–Russian collaboration.
What was measured: Cytogenetic measures of chromatin condensation in the cultured cells.
Where to read it: PubMed

A cytogenetic laboratory study on donor cells, not a study of the donors themselves.

Cell culture and in-vitro assays · 2011

Where labeled short peptides end up in cells, and how they bind DNA in vitro

Fedoreyeva LI, Kireev II, Khavinson VKh, Vanyushin BF. Biochemistry (Moscow). 2011;76:1210–1219.

What was studied: Fluorescently labeled short peptides: the tetrapeptide AEDG, the tripeptide EDR, and the tetrapeptide KEDG.
Model and setting: HeLa cells (an established human cell line) for localization; separate cell-free assays with DNA and oligonucleotides for binding.
What was measured: Fluorescence location within the cells; peptide–DNA interaction in the cell-free assays.
Where to read it: Publisher / DOI

Localization in a cell line is a laboratory observation about that model; the paper does not study oral intake.

Analytical chemistry · 2017

Is the synthetic sequence AEDG present in the natural pineal extract?

Khavinson VKh, Linkova NS, Tarnovskaya SI, et al. Bulletin of Experimental Biology and Medicine. 2017;164:41–43.

What was studied: The natural low-molecular-weight polypeptide complex of the pineal gland, analyzed for the presence of the tetrapeptide AEDG.
Model and setting: Analytical laboratory work (chromatography and mass spectrometry) on the extract itself; no cells, animals, or people.
What was measured: Whether, and in what proportion, the AEDG sequence is detectable in the natural complex.
Where to read it: Publisher / DOI · PubMed

Directly relevant to the natural-versus-synthetic distinction explained in the pillar article: it asks how the two kinds of material relate chemically.

Cell culture · 2019

AEDG, telomere length, and mitotic index in cultured human lymphocytes

Khavinson VKh, Pendina AA, Efimova OA, Tikhonov AV, Koltsova AS, Krapivin MI, Petrovskaia-Kaminskaia AV, Petrova LI, Lin'kova NS, Baranov VS. Bulletin of Experimental Biology and Medicine. 2019;168:123–126.

What was studied: The tetrapeptide AEDG, added to cell cultures.
Model and setting: Human blood lymphocytes induced to divide in culture (PHA-stimulated); a collaboration with a St. Petersburg genetics laboratory.
What was measured: Telomere length and mitotic index in the cultured cells.
Where to read it: Publisher / DOI · PubMed

A cytogenetic cell-culture study; complements the 2003 fibroblast experiment with a different cell type and method.

Cell culture · 2020

Short peptides and long-term culture of human oral stem cells

Sinjari B, Diomede F, Khavinson V, Mironova E, Linkova N, Trofimova S, Trubiani O, Caputi S. Stem Cell Reviews and Reports. 2020;16:159–166.

What was studied: Short synthetic peptides of the program, added to stem-cell cultures.
Model and setting: Human gingival mesenchymal stem cells kept in long-term culture; a collaboration with the University of Chieti–Pescara (Italy).
What was measured: Markers of cellular senescence and proliferation across culture passages.
Where to read it: Publisher / DOI

One of several papers from an Italian–Russian collaboration on stem-cell cultures.

Cell culture · 2020

AEDG and neuronal-differentiation markers in stem-cell culture

Khavinson V, Diomede F, Mironova E, Linkova N, Trofimova S, Trubiani O, Caputi S, Sinjari B. Molecules. 2020;25(3):609.

What was studied: The tetrapeptide AEDG, added to stem-cell cultures.
Model and setting: Human gingival mesenchymal stem cells induced toward neuronal differentiation in culture; the same Italian–Russian collaboration.
What was measured: Expression of genes and proteins used as markers of neuronal differentiation.
Where to read it: Publisher / DOI (open access) · PubMed

Open access; the full methods are available.

Cell culture · 2025 (with correction)

AEDG in several established human cell lines, from a group outside the program

Al-dulaimi S, Thomas R, Matta S, Roberts T. Biogerontology. 2025;26:178. Correction: Biogerontology. 2026;27:1 (published November 15, 2025).

What was studied: The synthetic tetrapeptide AEDG, supplied to the authors as a research material.
Model and setting: Normal human fibroblast and epithelial cell lines and several immortalized human cell lines, in culture.
What was measured: Telomere length, telomerase activity, and alternative lengthening of telomeres (ALT) in each cell type.
Where to read it: Corrected article / DOI · Correction notice

One of the few papers from a group outside the St. Petersburg program. Read the article together with its correction, which replaced Figures 1–3; it is a correction, not a retraction.

Molecular modeling

Three papers use computer simulation to ask what is structurally possible: how a short peptide might sit in the groove of DNA, or whether it could pass through the transporter proteins that carry amino acids across cell membranes. Modeling proposes hypotheses for experiments to test; it does not measure anything in a living system.

Molecular modeling · 2016

Docking models of nineteen short peptides with DNA

Khavinson VKh, Linkova NS, Tarnovskaya SI. Bulletin of Experimental Biology and Medicine. 2016;162:288–292.

What was studied: Nineteen short synthetic peptides from the program, modeled computationally.
Model and setting: Computer docking simulations; the introduction also reviews earlier cell and animal experiments by the group.
What was measured: Predicted binding configurations between each peptide and DNA sequences.
Where to read it: PubMed · Full text PDF (author-hosted)

A modeling paper proposes possible interactions; it is not an experiment on cells or people.

Molecular modeling · 2019

A systematic search for how short peptides could bind double-stranded DNA

Kolchina N, Khavinson V, Linkova N, Yakimov A, Baitin D, Afanasyeva A, Petukhov M. Nucleic Acids Research. 2019;47(20):10553–10563.

What was studied: Short peptides in general, screened computationally for DNA-binding motifs.
Model and setting: Molecular modeling and computational screening; published in a major international molecular-biology journal.
What was measured: Structural motifs predicted to allow peptide binding in the grooves of the DNA double helix.
Where to read it: Publisher (open access) · PubMed

Theoretical work; it maps what is structurally possible rather than what happens in a cell or an organism.

Molecular modeling · 2023

Could ultrashort peptides use the body's amino-acid transporters?

Khavinson VKh, Linkova NS, Rudskoy AI, Petukhov MG. Biomolecules. 2023;13(3):552.

What was studied: Twenty-six ultrashort peptides (two to three amino acids) from the program.
Model and setting: Computational docking against the structures of the LAT1, LAT2, and PEPT1 transporter proteins.
What was measured: Predicted binding to each transporter, compared with the transporters' known substrates.
Where to read it: Full text PDF (author-hosted) · Publisher / DOI

Addresses a question readers often ask — how a swallowed peptide could cross a membrane at all — as a modeling exercise, not as a measurement in people.

Review literature

Four reviews, three of them written by researchers from the program about their own work and one a systematic review with a stated search method. Reviews are the fastest way to find primary papers; they are not additional independent studies.

Review · 2009

Thirty-five years of the program, summarized by its authors

Khavinson VKh, Anisimov VN. Advances in Gerontology (Uspekhi Gerontologii). 2009;22(1):11–23. In Russian; English abstract on PubMed.

What was studied: The program's tissue extracts and synthetic peptides.
Model and setting: A narrative review of experimental and clinical work by the originating group up to 2009.
What was measured: Not an experiment.
Where to read it: PubMed

Russian-language; useful mainly for its reference list.

Review · 2010

The program's English-language overview

Anisimov VN, Khavinson VKh. Biogerontology. 2010;11:139–149.

What was studied: Multiple preparations from the program — tissue extracts and synthetic short peptides.
Model and setting: A narrative review summarizing the group's own animal experiments and clinical applications up to 2010.
What was measured: Not an experiment: it collects and discusses earlier publications.
Where to read it: PubMed · Publisher / DOI

The most-cited English-language overview. Written by researchers from the program it reviews; a review is not an additional independent study.

Review · 2013

The program's clinical reports, reviewed by its authors

Khavinson VKh, Kuznik BI, Ryzhak GA. Advances in Gerontology (Uspekhi Gerontologii). 2013;26(1):20–37. In Russian; English abstract on PubMed.

What was studied: The injectable preparations and later products of the program.
Model and setting: A narrative review of clinical reports, mostly Russian-language, up to 2013.
What was measured: Not an experiment.
Where to read it: PubMed

Companion to a first report on experimental studies. Russian-language; the abstract is in English.

Systematic review · 2021

A systematic review of the literature on short peptides and gene expression

Khavinson VKh, Popovich IG, Linkova NS, Mironova ES, Ilina AR. Molecules. 2021;26(22):7053.

What was studied: Short peptides studied in relation to gene expression, across the literature.
Model and setting: A systematic review with a stated search method; open access.
What was measured: Not an experiment: it collects and classifies published studies.
Where to read it: Publisher (open access) · PubMed Central

The most recent overview and the only one with a systematic search method; written by authors from the program.

Books and monographs

Two book-length works by the program's authors are frequently cited and worth knowing by name. They are not freely available online, so we list them without links.

Khavinson VKh. Peptides and Ageing. Neuroendocrinology Letters. 2002;23 (Supplement 3). A monograph-length review of the program's first three decades.
Khavinson VKh, Malinin VV. Gerontological Aspects of Genome Peptide Regulation. Basel: Karger; 2005. A book presenting the group's hypothesis of how short peptides interact with DNA.

Names that are easy to confuse

Several substances in these papers have similar names. Matching the exact name to the exact paper is the single most useful habit when reading this literature.

Similar names, different substances
Name	What it is	Appears in
Epithalamin / Epithalamine	A pineal-gland extract preparation from the original pharmaceutical work; given by injection.	2003 follow-up; 2006 and 2011 cohort reports
AEDG (also called Epitalon or Epithalon in the literature)	The synthetic tetrapeptide Ala-Glu-Asp-Gly, designed from the composition of the pineal extract. On this page it is referred to by sequence.	All animal studies; most laboratory studies
Thymalin	A thymus extract preparation from the original pharmaceutical work; given by injection.	2003 follow-up report
EDR, KEDG	The synthetic tripeptide Glu-Asp-Arg and tetrapeptide Lys-Glu-Asp-Gly, referred to here by sequence.	2011 localization study

What this library does not do

It does not summarize results. We describe what each study examined and measured; the authors' findings, effect sizes, and limitations are in the papers, and the links take you to them.
It does not display article titles. References are given in author–journal–year–pages form, the convention used by several physics and chemistry journals, so that this page describes the studies in its own neutral wording; each title and abstract is one click away on the linked page.
It is not systematic. These 21 papers and two books were selected as the most frequently cited entry points into the program. They are not the whole literature — the literature on AEDG alone runs to several dozen papers — and we have not searched for every favorable, null, or adverse report.
It does not rank evidence. No grading scheme has been applied. The labels (human, animal, laboratory, modeling, review) describe study type, not quality.
It does not concern retail products. The materials studied are the ones named in each paper. Whether any of them corresponds to a product sold today is a question for that product's label and documentation, not for this page.
It notes corrections where we found them. One paper (2025) has a published correction; read the two together. We did not run an exhaustive retraction check across all databases.

Questions about the research

Are these studies about the products sold today?

No. Each paper studies the material it names — Thymalin, Epithalamin, AEDG, or another specific synthetic sequence — under the conditions it describes. This page does not connect any of them to a retail product.

Why are there no results on this page?

Because a result without its methods, population, comparison group, and limitations is easy to misread, and because we would rather send you to the source than paraphrase it. Each entry tells you what was measured so you know what to look for when you open the paper.

Why do you write “AEDG” instead of “Epitalon”?

Epitalon (also spelled Epithalon) is a trade-style name for the synthetic tetrapeptide Ala-Glu-Asp-Gly. The sequence is unambiguous and is how the peptide is identified in the analytical and modeling papers, so we use it throughout. The names table above lists the equivalents.

Where can I read the full texts?

Several entries are open access: the 2019 Nucleic Acids Research paper, the 2020 Molecules paper, the 2021 systematic review, the 2016 and 2023 modeling papers (author-hosted PDFs), and the 2025 study with its correction. The rest link to publisher abstracts. The two books are in print and in university libraries.

Who wrote these papers?

Most come from Vladimir Khavinson's group and its collaborators at the St. Petersburg Institute of Bioregulation and Gerontology, sometimes with co-authors in Italy, Israel, Georgia, and Ukraine. One 2025 cell-line study comes from a group outside that program. Authorship and funding are stated in each paper and are worth reading before the results.

Is any of this research about taking peptides by mouth?

Not in the papers listed here. The human studies used injectable preparations; the animal studies used injections or added the peptide to the animals' medium; the laboratory studies added peptides directly to cells or modeled them on a computer. The 2023 modeling paper asks, computationally, whether ultrashort peptides could use intestinal transporters — a hypothesis, not a measurement.

How often is this page updated?

We re-check the links and look for new corrections when we revise the page; the date at the top is the last revision. If you know of a correction or retraction we have missed, please let us know.

Full reference list

All 21 publications in the order they appear above, in author–journal–year–pages form, each with its links. The article title and abstract are on the linked page.

Khavinson VKh, Morozov VG. Neuro Endocrinology Letters. 2003;24(3–4):233–240. — Publisher abstract · PubMed
Korkushko OV, Khavinson VKh, Shatilo VB, Antonyuk-Shcheglova IA. Bulletin of Experimental Biology and Medicine. 2006;142:356–359. — Publisher / DOI
Korkushko OV, Khavinson VKh, Shatilo VB, Antonyk-Sheglova IA. Bulletin of Experimental Biology and Medicine. 2011;151:366–369. — Publisher / DOI
Khavinson VKh, Izmaylov DM, Obukhova LK, Malinin VV. Mechanisms of Ageing and Development. 2000;120:141–149. — Publisher / DOI · PubMed
Khavinson V, Goncharova N, Lapin B. Neuro Endocrinology Letters. 2001;22(4):251–254. — PubMed
Kossoy G, Zandbank J, Tendler E, Anisimov V, Khavinson V, Popovich I, Zabezhinski M, Zusman I, Ben-Hur H. Journal of Endocrinological Investigation. 2003. — Publisher / DOI · PubMed
Khavinson VKh, Bondarev IE, Butyugov AA. Bulletin of Experimental Biology and Medicine. 2003;135:590–592. — Publisher / DOI · PubMed
Khavinson VKh, Lezhava TA, Monaselidze JR, Jokhadze TA, Dvalishvili NA, Bablishvili NK, Trofimova SV. Neuro Endocrinology Letters. 2003;24(5):329–333. — PubMed
Fedoreyeva LI, Kireev II, Khavinson VKh, Vanyushin BF. Biochemistry (Moscow). 2011;76:1210–1219. — Publisher / DOI
Khavinson VKh, Linkova NS, Tarnovskaya SI, et al. Bulletin of Experimental Biology and Medicine. 2017;164:41–43. — Publisher / DOI · PubMed
Khavinson VKh, Pendina AA, Efimova OA, Tikhonov AV, Koltsova AS, Krapivin MI, Petrovskaia-Kaminskaia AV, Petrova LI, Lin'kova NS, Baranov VS. Bulletin of Experimental Biology and Medicine. 2019;168:123–126. — Publisher / DOI · PubMed
Sinjari B, Diomede F, Khavinson V, Mironova E, Linkova N, Trofimova S, Trubiani O, Caputi S. Stem Cell Reviews and Reports. 2020;16:159–166. — Publisher / DOI
Khavinson V, Diomede F, Mironova E, Linkova N, Trofimova S, Trubiani O, Caputi S, Sinjari B. Molecules. 2020;25(3):609. — Publisher / DOI (open access) · PubMed
Al-dulaimi S, Thomas R, Matta S, Roberts T. Biogerontology. 2025;26:178. Correction: Biogerontology. 2026;27:1 (published November 15, 2025). — Corrected article / DOI · Correction notice
Khavinson VKh, Linkova NS, Tarnovskaya SI. Bulletin of Experimental Biology and Medicine. 2016;162:288–292. — PubMed · Full text PDF (author-hosted)
Kolchina N, Khavinson V, Linkova N, Yakimov A, Baitin D, Afanasyeva A, Petukhov M. Nucleic Acids Research. 2019;47(20):10553–10563. — Publisher (open access) · PubMed
Khavinson VKh, Linkova NS, Rudskoy AI, Petukhov MG. Biomolecules. 2023;13(3):552. — Full text PDF (author-hosted) · Publisher / DOI
Khavinson VKh, Anisimov VN. Advances in Gerontology (Uspekhi Gerontologii). 2009;22(1):11–23. In Russian; English abstract on PubMed. — PubMed
Anisimov VN, Khavinson VKh. Biogerontology. 2010;11:139–149. — PubMed · Publisher / DOI
Khavinson VKh, Kuznik BI, Ryzhak GA. Advances in Gerontology (Uspekhi Gerontologii). 2013;26(1):20–37. In Russian; English abstract on PubMed. — PubMed
Khavinson VKh, Popovich IG, Linkova NS, Mironova ES, Ilina AR. Molecules. 2021;26(22):7053. — Publisher (open access) · PubMed Central

About this page. Published by Vita Stream Inc for educational purposes. It is a selected bibliography with study-design notes. It does not summarize findings, does not evaluate the clinical effectiveness of any product, and is not medical advice. Discuss questions about your own health with a qualified healthcare professional.

Cite this library: Vita Stream Inc. Peptide Bioregulator Research Library: Publications from the Khavinson Program. Updated October 1, 2026. Permanent URL.